State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong 266237, China; Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, Shandong 266237, China. Electronic address: lishengying@sdu.edu.cn. State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong 266237, China; Shandong Provincial Key Laboratory of Synthetic Biology, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, Shandong 266101, China. Department of Biochemistry, Campus B2.2, Saarland University, D-66123 Saarbrücken, Germany. Electronic address: ritabern@mx.uni-saarland.de.